Most botanical references place the genus Mimulus in the Snapdragon family, Scrophulariaceae; but modern genetic studies have persuaded botanists to remove it to the family Phrymaceae, the Lopseed family, which had previously had only one species in it.
Gray’s description of this species depends on his description of M. ringens, so we print that description in brackets:
MÍMULUS L. MONKEY FLOWER. Calyx prismatic, 5-angled, 5-toothed, the uppermost tooth largest. Upper lip of corolla erect or reflexed-spreading, 2-lobed; lower spreading, 3-lobed. Stigma 2-lobed; lobes ovate. Seeds numerous. — Herbs, with opposite (rarely whorled) leaves, and mostly handsome flowers. (Diminutive of mimus, a buffoon, from the grinning corolla.)
Corolla violet-purple (rarely white); erect glabrous perennials; leaves feather-veined.
[M. ríngens L. Stem square, 1 m. or less high; leaves oblong or lanceolate, pointed, clasping by a heart-shaped base, serrate; peduncles longer than the flower; calyx-teeth taper-pointed, nearly equal; corolla personate, 2-4 cm. long. — Wet places, N. B. to Man., and southw. June-Sept.]
M. alàtus Ait. Stem winged at the angles; leaves oblong-ovate, tapering into a petiole; peduncles shorter than the very short-toothed calyx; otherwise like the preceding. — Wet places, Ct. to s. Ont., Kan., and southw.
Adaptable to many different lighting conditions; we wrote earlier that Foxglove Beardtongue “likes a sunny open field or clearing, although it will tolerate some shade,” but this one was growing in quite deep shade in a thicket in Schenley Park. It was blooming in early June.
A relative of snapdragons and Butter-and-Eggs, this cheery flower also bears a passing resemblance to a foxglove, whence both the common and scientific names. The name “Beardtongue” comes from the hairy stamen visible in each flower.
Most earlier botanical references spell the genus name Pentstemon, which may be more etymologically correct but apparently is not the way it was spelled in the original description.
Traditionally, botanists placed snapdragons and their allies in the Snapdragon or Figwort family, Scrophulariaceae; but modern genetic research has led botanists to move them into the Plantain family, Plantaginaceae.
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Gray makes this species a variety of P. laevigatus, so we turn to Alphonso Wood for a description of the genus and the species more in line with the consensus of modern botanists:
PENTSTEMON, L. Beard-tongue. Calyx deeply 5-cleft. Cor. elongated, often ventricous, lower lip 3-lobed, spreading. The fifth filament (tongue) sterile, bearded, longer than the rest or about as long; anth. smooth. Seeds numerous, angular, not margined. Perennial N. American, branching, paniculate. Leaves opposite, the lower petiolate, upper sessile or clasping. Flowers showy, red, violet, blue, or white, in Summer.
Native E. of the Mississippi River, sometimes cultivated.
Leaves undivided, serrulate. Sterile filament (tongue) bearded.
P. digitalis N. Glabrous; leaves elliptic to lanceolate, the upper clasping; flowers many, large, corolla tube abruptly enlarged to bell-form, pale blue or purplish, 12—15″ long, throat widely open, beardless. Rich soils, Pa., W. and S.
These intensely blue flowers are so tiny that we often overlook them in our lawns, but they are one of the first cheerful signs of spring. They are alien invaders, and perhaps they have caused untold damage to our environment; but it’s hard to be angry at a plant that’s both tiny and beautiful. This plant was one of a patch growing in a lawn in Beechview, where it was blooming at the end of March.
The flowers of the Persian Speedwell have yellow centers, fading to white veined with blue, the blue predominating toward the outer edges of the petals, and giving the overall impression of a blue flower from a short distance. The leaves are roundish, sessile near the top of thestem and on short petioles below, gently toothed, somewhat hairy. The plant seldom exceeds the height of a few inches, and can often pass unmolested under a lawnmower blade.
This description will have to do, since Gray and his contemporaries did not describe the plant. Between their time and ours it has spread to every state in the union except Hawaii and North Dakota.